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The painting’s brilliant range of tones has now been revealed following careful restoration. An unusual Botticelli work for having been painted on canvas and not wood.

One of the world’s most famous sculptures, the sisterly trio of nudes was commissioned by the Duke of Bedford in 1815. Canova has all but turned marble into living flesh.

Velazquez’ creation of mood through strong contrast was unprecedented in Spain when he produced this work in 1618.

One of the most celebrated paintings by a Scottish painter. The fun-loving minister depicted is believed to have been a member of the prestigious Edinburgh Skating Club.

The seven works depicting the rites of Christianity evoke grand theatricality; they are considered the finest pieces by Nicolas Pousssin, founder of French Classical painting.

Works by Leonardo da Vinci and Raphael stand out. Leonardo’s drawing of Studies of a Dog’s Paw shows his ability to convey a world in a few strokes, while Raphael’s Bridgewater Madonna is alive with tenderness.

Look for the superb portraits by Ramsay, Raeburn and Guthrie, and Pitlessie Fair painted at the age of 16 by Sir David Wilkie. Autumn in Glen cairn is an outstanding work by James Paterson.

The pick of the best must include Rembrandt’s world-weary Self-Portrait aged 51 , Gainsborough’s 1777 hit at the Royal Academy The Honourable Mrs Graham , and Van Dyck’s The Lomellini Family , complete with a capering dog. Another favourite is Wilkie’s The Greenwich Pensioner .

Of the 5 Monets here, A Seascape, Shipping by Moonlight is a rare palette knife and brush application, while Poplars on the Epte is vintage Monet. Degas’ A Group of Dancers also features.

William Playfair chose a Neoclassical style to link Old and New Edinburgh. He also built the neighbouring Royal Scottish Academy and the two buildings have recently been joined.